THE EVOLUTIONARY STRATEGIES THAT SHAPE ECOSYSTEMS In 1837 a young Charles Darwin took his notebook, wrote “I think”, and then sketched a rudimentary, stick-like tree. Each branch of Darwin’s tree of life told a story of survival and adaptation – adaptation of animals and plants not just to the environment but also to life with other living things. However, more than 150 years since Darwin published his singular idea of natural selection, the science of ecology has yet to account for how contrasting evolutionary outcomes affect the ability of organisms to coexist in communities and to regulate ecosystem functioning. In this book Philip Grime and Simon Pierce explain how evidence from across the world is revealing that, beneath the wealth of apparently limitless and bewildering variation in detailed structure and functioning, the essential biology of all organisms is subject to the same set of basic interacting constraints on life-history and physiology. The inescapable resulting predicament during the evolution of every species is that, according to habitat, each must adopt a predictable compromise with regard to how they use the resources at their disposal in order to survive. The compromise involves the investment of resources in either the effort to acquire more resources, the tolerance of factors that reduce metabolic performance, or reproduction. This three-way trade-off is the irreducible core of the universal adaptive strategy theory which Grime and Pierce use to investigate how two environmental filters selecting, respectively, for convergence and divergence in organism function determine the identity of organisms in communities, and ultimately how different evolutionary strategies affect the functioning of ecosystems. This book refl ects an historic phase in which evolutionary processes are finally moving centre stage in the effort to unify ecological theory, and animal, plant and microbial ecology have begun to find a common theoretical framework. Companion website This book has a companion website www.wiley.com/go/grime/evolutionarystrategies with Figures and Tables from the book for downloading.

As a novel endeavour in ecological science, this book focuses on amajor issue in organismal life on Earth:species coexistence. Thebook crosses the usual disciplinary boundaries betweenpalaeobiology, ecology and evolutionary biology and provides atimely overview of the patterns and processes of species diversityand coexistence on a range of spatio-temporal scales. In thisunique synthesis, the author offers a critical and penetratingexamination of the concepts and models of coexistence and communitystructure, thus making a valuable contribution to the field ofcommunity ecology. There is an emphasis on clarity andaccessibility without sacrificing scientific rigour, making thisbook suitable for both advanced students and individual researchersin ecology, palaeobiology and environmental and evolutionarybiology. Comprehensive and contemporary synthesis. Pulls together the aggregate influence of evolution and ecologyon patterns in communities. Balanced mix of theory and empirical work. Clearly structured chapters with short introduction andsummary.

This book provides an up-to-date coverage of green (vegetated) roof research, design, and management from an ecosystem perspective. It reviews, explains, and poses questions about monitoring, substrate, living components and the abiotic, biotic and cultural aspects connecting green roofs to the fields of community, landscape and urban ecology. The work contains examples of green roof venues that demonstrate the focus, level of detail, and techniques needed to understand the structure, function, and impact of these novel ecosystems. Representing a seminal compilation of research and technical knowledge about green roof ecology and how functional attributes can be enhanced, it delves to explore the next wave of evolution in green technology and defines potential paths for technological advancement and research.

How do plants make a living? Some plants are gamblers, others are swindlers. Some plants are habitual spenders while others are strugglers and miserly savers. Plants have evolved a spectacular array of solutions to the existential problems of survival and reproduction in a world where resources are scarce, disturbances can be deadly, and competition is cut-throat. Few topics have both captured the imagination and furrowed the brows of plant ecologists, yet no topic is more important for understanding the assembly of plant communities, predicting plant responses to global change, and enhancing the restoration of our rapidly degrading biosphere. The vast array of plant strategy models that characterize the discipline now require synthesis. These models tend to emphasize either life history strategies based on demography, or functional strategies based on ecophysiology. Indeed, this disciplinary divide between demography and physiology runs deep and continues to this today. The goal of this accessible book is to articulate a coherent framework that unifies life history theory with comparative functional ecology to advance prediction in plant ecology. Armed with a deeper understanding of the dimensionality of life history and functional traits, we are now equipped to quantitively link phenotypes to population growth rates across gradients of resource availability and disturbance regimes. Predicting how species respond to global change is perhaps the most important challenge of our time. A robust framework for plant strategy theory will advance this research agenda by testing the generality of traits for predicting population dynamics.

Biological diversity, the variety of living organisms on Earth, is traditionally viewed as the diversity of taxa, and species in particular. However, other facets of diversity also need to be considered for a comprehensive understanding of evolutionary and ecological processes. This novel book demonstrates the advantages of adopting a functional approach to diversity in order to improve our understanding of the functioning of ecological systems and theircomponents. The focus is on plants, which are major components of these systems, and for which the functional approach has led to major scientific advances over the last 20 years. PlantFunctional Diversity presents the rationale for a trait-based approach to functional diversity in the context of comparative plant ecology and agroecology. It demonstrates how this approach can be used to address a number of highly debated questions in plant ecology pertaining to plant responses to their environment, controls on plant community structure, ecosystem properties, and the services these deliver to human societies. This research level text will be of particular relevance and use tograduate students and professional researchers in plant ecology, agricultural sciences and conservation biology.

Life histories can be defined as the means by which individuals (or more precisely genotypes) vary their age- or stage-specific expenditures of reproductive effort in response to genetic, phenotypic, and environmental correlates of survival and fecundity. Life histories reflect the expression of traits most closely related to individual fitness, such as age and size at maturity, number and size of offspring, and the timing of the expression of those traits throughout an individual's life. In addition to addressing questions of fundamental importance to ecology and evolution, life-history research plays an integral role in species conservation and management. This accessible primer encompasses the basic concepts, theories, and applied elements of life history evolution, including patterns of trait variability, underlying mechanisms of plastic/evolutionary change, and the practical utility of life-history traits as metrics of species/population recovery, sustainable exploitation, and risk of extinction. Empirical examples are drawn from the entire spectrum of life. A Primer of Life Histories is designed for readers from a broad range of academic backgrounds and experience including graduate students and researchers of ecology and evolutionary biology. It will also be useful to a more applied audience of academic/government researchers in fields such as wildlife biology, conservation biology, fisheries science, and the environmental sciences.

The novelty of the book is a strong focus on perception, perspectives and prediction by scientists with profound insight into the ecology of ecosystems or into human demands and activity. The challenge is to bridge from empirical data and the knowledge of the past to the possibilities of the performance in the future. We assume that there is scope for more cooperation between the fields of ecology and practical philosophy or other social sciences in organising ecosystems and shaping the cultural future of humankind, and that such collaboration should be accorded considerably more priority. This book deals with environmental processes seen within a framework of the nature of ecosystems and human cultures. The future of the environment, the development of ecosystems and effective nature conservation management are the essentials of this book. Human nature and culture, and in particular their interactions, are interpreted as a set of rules and as given. The aim is not only to assess the significance of human influence on species composition and biodiversity but also to weigh up the subsequent potentials for action. In this book we will analyze the problems independently of one another, even if they are interconnected. This book focuses on perspectives and prognoses for the impacts of anthropogenic activity on ecosystems and thus on species conservation. Its goal is to improve assessments of the impacts of human activity on the environment. We are aware that prognoses have very often proven to be false. It is difficult to impossible to be able to predict with precision how evolution and ecosystems will change in future under anthropogenic influence. This strengthens our resolve to attempt to retain the highest possible degree of scientific integrity and professionalism and not to shy away from expressing the uncertainty of our own ideas and prognoses. We venture prognoses in this book and we will fail. However, we hope that we will be wrong on the right side.