This classic text, first published in 1935, is once again available. Still the standard reference in the English language, Principles of Insect Morphology is considered the author's masterpiece. A talented artist as well as one of the leading entomologists of his day, Robert E. Snodgrass produced a wealth of publications that display an accuracy and precision still unsurpassed. The 19 chapters in this volume cover each group of insect organs and their associated structures, at the same time providing a coherent morphological view of their fundamental nature and apparent evolution. To accomplish this aim, Snodgrass compares insect organs with those of other arthropods. Each chapter concludes with a glossary of terms. The 319 multipart illustrations are an invaluable source of information and have never been duplicated. This edition includes a new foreword by George Eickwort, Professor of Entomology at Cornell University, which relates the book to today's courses in insect morphology. Republication of this textbook will provide another generation of students with an essential foundation for their studies in entomology.

seem as appropriate now as the original balance was when Dr A. D. Imms' textbook was first published over fifty years ago. There are 35 new figures, all based on published illustrations, the sources of which are acknowledged in the captions. We are grateful to the authors concerned and also to Miss K. Priest of Messrs Chapman & Hall, who saved us from many errors and omissions, and to Mrs R. G. Davies for substantial help in preparing the bibliographies and checking references. London O.W.R. May 1976 R.G.D. Part I ANATOMY AND PHYSIOLOGY Chapter I INTRODUCTION Definition of the Insecta (Hexapoda) The insects are tracheate arthropods in which the body is divided into head, thorax and abdomen. A single pair of antennae (homologous with the anten nules of the Crustacea) is present and the head also bears a pair of mandibles and two pairs of maxillae, the second pair fused medially to form the labium. The thorax carries three pairs of legs and usually one or two pairs of wings. The abdomen is devoid of ambulatory appendages, and the genital opening is situated near the posterior end of the body. Postembryonic development is rarely direct and a metamorphosis usually occurs.

In the last decades a remarkable renaissance has materialized in insect morphology, mainly triggered by the development of new cutting-edge technologies. This is an exciting time for biological synthesis where the mysteries and data derived from genomes can be combined with centuries of data from morphology and development. And, now, more than ever, detailed knowledge of morphology is essential to understanding the evolution of all groups of organisms. In this “age of phylogenomics” researchers rely on morphological data to support molecular findings, test complex evolutionary scenarios, and for placing fossil taxa. This textbook provides an in-depth treatment of the structures and the phylogeny of the megadiverse Hexapoda. The first part presents an up-to-date overview of general insect morphology with detailed drawings, scanning electron micrographs, and 3-D reconstructions. Also included is a chapter covering innovative morphological techniques (e.g., μ-computer tomography, 3-D modeling), brief treatments of insect development and phylogenetic methods, and a comprehensive morphological glossary. The second part is of a modern synthesis of insect systematics that includes taxon-specific morphological information for all Orders. The work is an invaluable reference for students and researchers working in all facets of biology and is a must for evolutionary biologists. A detailed understanding of morphology is essential in unraveling phylogenetic relationships and developing complex evolutionary scenarios. Increasingly researchers in phylogenomics are re/turning to morphological data to support their findings, while the development of new cutting-edge technologies has further increased interest in this growing field. This definitive handbook provides an in-depth treatment of insect morphology. The first part presents an up-to-date overview of insect morphology with detailed drawings, brilliant scanning electron micrographs and 3-D reconstructions as interactive PDFs. This is complemented by a chapter on innovative morphological techniques (e.g., μ-computer tomography, 3-D modeling) and a comprehensive morphological glossary. The second part treats the state of the art in insect systematics and includes taxon-specific morphological information for all orders. Systematics are treated formally, with for example the arguments for relationships (“apomorphies”) always listed explicitly. The work is a useful reference for students and researchers working in different fields of biology and a must for those dealing with insects from an evolutionary perspective.

The arthropods contain more species than any other animal group, but the evolutionary pathways which led to their current diversity are still an issue of controversy. Arthropod Relationships provides an overview of our current understanding, responding to the new data arising from sequencing DNA, the discovery of new Cambrian fossils as direct evidence of early arthropod history, and developmental genetics. These new areas of research have stimulated a reconsideration of classical morphology and embryology. Arthropod Relationships is the first synthesis of the current debate to emerge: not since the volume edited by Gupta was published in 1979 has the arthropod phylogeny debate been, considered in this depth and breadth. Leaders in the various branches of arthropod biology have contributed to this volume. Chapters focus progressively from the general issues to the specific problems involving particular groups, and thence to a consideration of embryology and genetics. This wide range of disciplines is drawn on to approach an understanding of arthropod relationships, and to provide the most timely account of arthropod phylogeny. This book should be read by evolutionary biologists, palaeontologists, developmental geneticists and invertebrate zoologists. It will have a special interest for post-graduate students working in these fields.

Insects are the most diverse group of organisms in the 3 billion-year history of life on Earth, and the most ecologically dominant animals on land. This book chronicles for the first time the complete evolutionary history of insects: their living diversity, relationships and 400 million years of fossils. Whereas other volumes have focused on either living species or fossils, this is the first comprehensive synthesis of all aspects of insect evolution. The book is illustrated with 955 photo- and electronmicrographs, drawings, diagrams, and field photos, many in full colour and virtually all of them original. The book will appeal to anyone engaged with insect diversity: professional entomologists and students, insect and fossil collectors, and naturalists.

The renowned German reference work The Handbuch der Zoologie/Handbook of Zoology was founded in the 1920's by Professor Willi Kükenthal in Berlin and treated the complete animal kingdom from single cell organisms to mammals in eight thematic volumes: Volume I Protozoa, Porifera, Colenteratea, Mesozoa (1925); Volume II Worms (1933/34); Volume III Arthropoda ex. Insecta (1927/1932); Volume IV Arthopoda: Insecta; Volume V Solenogastres, Mollusca, Echinoderma (1925); Volume VI Pisces / Amphibia (1930); Volume VII Reptilia / Aves (1931); Volume VIII Mammalia. The Volumes Insecta (Eds. N.P. Kristensen, R.G. Beutel) and Mammalia (Eds. M.S. Fischer, H. Schliemann) continued publication into the present with the most recent contributions in English language. Covering nearly 100 years of zoological research, the Handbook of Zoology represents a vast store of knowledge. But with the speed of scientific discovery in the past decades, a new edition of the Handbook in a new form is required. Beginning in 2010 the Handbook of Zoology will be restructured and offered additionally as a database (Zoology Online) which can be easily searched and rapidly updated. The eight thematic volumes will be replaced with smaller and more flexible groupings that reflect the current state of phylogenetic knowledge. Faster publication times through online-prepublication, reference linking, forward linking and multimedia presentations will make the Handbook of Zoology highly attractive to both authors and users. Aims and Scope The Handbook of Zoology aims to provide an in depth treatment of the entire animal kingdom from the lower invertebrates to the mammals. It publishes comprehensive overviews on animal systematics and morphology as well as extensive coverage of physiology, behaviour, ecology and applied aspects of zoological research. Volumes in progress include Nemathelminthes and Gnathifera, Annelida, Mollusca, Arthropoda, Arthropoda: Insecta, and Mammalia. Although our knowledge regarding many taxonomic groups has grown enormously over the last decades, it is still the ambition of the Handbook to be comprehensive in the sense that text and references together provide a solid basis for further research. Editors and authors seek a balance between describing species richness and diversity, explaining the importance of certain groups in a phylogenetic context and presenting a review of available knowledge and up-to-date reference literature. New contributions to the series present the combined effort of an international team of editors and authors, entirely published in English and explicitly addressing the international scientific community.

More than two thirds of all living organisms described to date belong to the phylum Arthropoda. But their diversity, as measured in terms of species number, is also accompanied by an amazing disparity in terms of body form, developmental processes, and adaptations to every inhabitable place on Earth, from the deepest marine abysses to the earth surface and the air. The Arthropoda also include one of the most fashionable and extensively studied of all model organisms, the fruit-fly, whose name is not only linked forever to Mendelian and population genetics, but has more recently come back to centre stage as one of the most important and more extensively investigated models in developmental genetics. This approach has completely changed our appreciation of some of the most characteristic traits of arthropods as are the origin and evolution of segments, their regional and individual specialization, and the origin and evolution of the appendages. At approximately the same time as developmental genetics was eventually turning into the major agent in the birth of evolutionary developmental biology (evo-devo), molecular phylogenetics was challenging the traditional views on arthropod phylogeny, including the relationships among the four major groups: insects, crustaceans, myriapods, and chelicerates. In the meantime, palaeontology was revealing an amazing number of extinct forms that on the one side have contributed to a radical revisitation of arthropod phylogeny, but on the other have provided evidence of a previously unexpected disparity of arthropod and arthropod-like forms that often challenge a clear-cut delimitation of the phylum.