The educational benefits of replacing in-class lectures with hands-on activities are clear. Such active learning is a natural fit for paleontology, which can provide opportunities for examining fossils, analyzing data and writing. Additionally, there are a number of topics in the field that are exciting to geology majors and non-majors alike: very few can resist the lure of dinosaurs, huge meteor impacts, vicious Cretaceous sharks or a giant Pleistocene land mammal. However, it can seem difficult to introduce these techniques into a large general education class full of non-majors: paleontological specimens provide a natural starting point for hands-on classroom activities, but in a large class it is not always practical or possible to provide enough fossil material for all students. The Element introduces different types of active learning approaches, and then explains how they have been applied to a large introductory paleontology class for non-majors.

People hold a variety of prior conceptions that impact their learning. Prior conceptions that include erroneous or incomplete understandings represent a significant barrier to durable learning, as they are often difficult to change. While researchers have documented students' prior conceptions in many areas of geoscience, little is known about prior conceptions involving paleontology. In this Element, data on student prior conceptions from two introductory undergraduate paleontology courses are presented. In addition to more general misunderstandings about the nature of science, many students hold incorrect ideas about methods of historical geology, Earth history, ancient life, and evolution. Of special note are student perceptions of the limits of paleontology as scientific inquiry. By intentionally eliciting students' prior conceptions and implementing the pedagogical strategies described in other Elements in this series, lecturers can shape instruction to challenge this negative view of paleontology and improve student learning.

Hands-on learning in paleontology, and geology in general, is fairly common practice. Students regularly use rocks, fossils, and data in the classroom throughout their undergraduate career, but they typically do it sitting in a chair in a lab. Kinesthetic learning is a teaching model that requires students to be physically active while learning. Students may be involved in a physical activity during class or might be using their own bodies to model some important concept. This Element briefly discusses the theory behind kinesthetic learning and how it fits into a student-centered, active-learning classroom. It then describes in detail methods for incorporating it into student exercises on biostratigraphy, assessment of sampling completeness, and modeling evolutionary processes. Assessment data demonstrates that these exercises have led to significantly improved student learning outcomes tied to these concepts.

New online resources are opening doors for education and outreach in the Earth sciences. One of the most innovative online earth science portals is Macrostrat and its mobile client Rockd - an interface that combines geolocated geological maps with stratigraphic information, lithological data, and crowd-sourced images and descriptions of outcrops. These tools provide a unique educational opportunity for students to interact with primary geological data, create connections between local outcrops and global patterns, and make new field observations. Rockd incorporates an aspect of social media to its platform, which creates a sense of community for users. This Element outlines these resources, gives instructions on how to use them, and provides examples of how to integrate these resources into a variety of paleontology and earth science courses.

Lecturing has been a staple of university pedagogy, but a shift is ongoing because of evidence that active engagement with content helps strengthen learning and build more advanced skills. The flipped classroom, which delivers content to students outside of the class meeting, is one approach to maximize time for active learning. The fundamental benefit of a flipped class is that students learn more, but ensuring student preparation and engagement can be challenging. Evaluation policies can provide incentives to guide student effort. Flipping a class requires an initial time commitment, but the workload associated with evaluating student work during the course can be mitigated. The personal interactions from active learning are extremely rewarding for students and instructors, especially when class sizes are small and suitable room layouts are available. Overall, flipping a course doesn't require special training, just a willingness to experiment, reflect, and adjust.

The principles of stratigraphic paleobiology can be readily applied to the nonmarine fossil record. Consistent spatial and temporal patterns of accommodation and sedimentation in sedimentary basins are an important control on stratigraphic architecture. Temperature and precipitation covary with elevation, causing significant variation in community composition, and changes in base level cause elevation to undergo predictable changes. These principles lead to eight sets of hypotheses about the nonmarine fossil record. Three relate to long-term and cyclical patterns in the preservation of major fossil groups and their taphonomy, as well as the occurrence of fossil concentrations. The remaining hypotheses relate to the widespread occurrence of elevation-correlated gradients in community composition, long-term and cyclical trends in these communities, and the stratigraphic position of abrupt changes in community composition. Testing of these hypotheses makes the stratigraphic paleobiology of nonmarine systems a promising area of investigation.

Placing evolutionary events in the context of geological time is a fundamental goal in paleobiology and macroevolution. In this Element we describe the tripartite model used for Bayesian estimation of time calibrated phylogenetic trees. The model can be readily separated into its component models: the substitution model, the clock model and the tree model. We provide an overview of the most widely used models for each component and highlight the advantages of implementing the tripartite model within a Bayesian framework.