Professor Levins, one of the leading explorers in the field of integrated population biology, considers the mutual interpenetration and joint evolution of organism and environment, occurring on several levels at once. Physiological and behavioral adaptations to short-term fluctuations of the environment condition the responses of populations to long-term changes and geographic gradients. These in turn affect the way species divide the environments among themselves in communities, and, therefore, the numbers of species which can coexist. Environment is treated here abstractly as pattern: patchiness, variability, range, etc. Populations are studied in their patterns: local heterogeneity, geographic variability, faunistic diversity, etc.

Leading researchers in evolutionary developmental biology seek linkages between, and a synthesis of, development, physiology, endocrinology, ecology, and evolution. Evolutionary developmental biology, also known as evo-devo or EDB, seeks to find links between development and evolution by opening the "black box" of development's role in evolution and in the evolution of developmental mechanisms. In particular, this volume emphasizes the roles of the environment and of hormonal signaling in evo-devo. It brings together a group of leading researchers to analyze the dynamic interaction of environmental factors with developmental and physiological processes and to examine how environmental signals are translated into phenotypic change, from the molecular and cellular level to organisms and groups of organisms. Taken together, these chapters demonstrate the crucial roles of those processes of genetic, developmental, physiological, and hormonal change that underpin evolutionary change in development, morphology, physiology, behavior, and life-history. Part I investigates links between environmental signals and developmental processes that could be preserved over evolutionary time. Several contributors evaluate the work of the late Ryuichi Matsuda, especially his emphasis on the role of the external environment in genetic change and variability ("pan-environmentalism"). Other contributors in part I analyze different aspects of environmental-genetic-evolutionary linkages, including the importance of alternate ontogenies in evolution and the paradox of stability over long periods of evolutionary time. Part II examines the plasticity that characterizes much of development, with contributors discussing such topics as gene regulatory networks and heterochronicity. Part III analyzes the role of hormones and metamorphosis in the evolution of such organisms with alternate life-history stages as lampreys, amphibians, and insects.

The seemingly innocent observation that the activities of organisms bring about changes in environments is so obvious that it seems an unlikely focus for a new line of thinking about evolution. Yet niche construction--as this process of organism-driven environmental modification is known--has hidden complexities. By transforming biotic and abiotic sources of natural selection in external environments, niche construction generates feedback in evolution on a scale hitherto underestimated--and in a manner that transforms the evolutionary dynamic. It also plays a critical role in ecology, supporting ecosystem engineering and influencing the flow of energy and nutrients through ecosystems. Despite this, niche construction has been given short shrift in theoretical biology, in part because it cannot be fully understood within the framework of standard evolutionary theory. Wedding evolution and ecology, this book extends evolutionary theory by formally including niche construction and ecological inheritance as additional evolutionary processes. The authors support their historic move with empirical data, theoretical population genetics, and conceptual models. They also describe new research methods capable of testing the theory. They demonstrate how their theory can resolve long-standing problems in ecology, particularly by advancing the sorely needed synthesis of ecology and evolution, and how it offers an evolutionary basis for the human sciences. Already hailed as a pioneering work by some of the world's most influential biologists, this is a rare, potentially field-changing contribution to the biological sciences.

Evolution: Components and Mechanisms introduces the many recent discoveries and insights that have added to the discipline of organic evolution, and combines them with the key topics needed to gain a fundamental understanding of the mechanisms of evolution. Each chapter covers an important topic or factor pertinent to a modern understanding of evolutionary theory, allowing easy access to particular topics for either study or review. Many chapters are cross-referenced. Modern evolutionary theory has expanded significantly within only the past two to three decades. In recent times the definition of a gene has evolved, the definition of organic evolution itself is in need of some modification, the number of known mechanisms of evolutionary change has increased dramatically, and the emphasis placed on opportunity and contingency has increased. This book synthesizes these changes and presents many of the novel topics in evolutionary theory in an accessible and thorough format. This book is an ideal, up-to-date resource for biologists, geneticists, evolutionary biologists, developmental biologists, and researchers in, as well as students and academics in these areas and professional scientists in many subfields of biology. - Discusses many of the mechanisms responsible for evolutionary change - Includes an appendix that provides a brief synopsis of these mechanisms with most discussed in greater detail in respective chapters - Aids readers in their organization and understanding of the material by addressing the basic concepts and topics surrounding organic evolution - Covers some topics not typically addressed, such as opportunity, contingency, symbiosis, and progress

The hominin fossil record documents a history of critical evolutionary events that have ultimately shaped and defined what it means to be human, including the origins of bipedalism; the emergence of our genus Homo; the first use of stone tools; increases in brain size; and the emergence of Homo sapiens, tools, and culture. The Earth's geological record suggests that some evolutionary events were coincident with substantial changes in African and Eurasian climate, raising the possibility that critical junctures in human evolution and behavioral development may have been affected by the environmental characteristics of the areas where hominins evolved. Understanding Climate's Change on Human Evolution explores the opportunities of using scientific research to improve our understanding of how climate may have helped shape our species. Improved climate records for specific regions will be required before it is possible to evaluate how critical resources for hominins, especially water and vegetation, would have been distributed on the landscape during key intervals of hominin history. Existing records contain substantial temporal gaps. The book's initiatives are presented in two major research themes: first, determining the impacts of climate change and climate variability on human evolution and dispersal; and second, integrating climate modeling, environmental records, and biotic responses. Understanding Climate's Change on Human Evolution suggests a new scientific program for international climate and human evolution studies that involve an exploration initiative to locate new fossil sites and to broaden the geographic and temporal sampling of the fossil and archeological record; a comprehensive and integrative scientific drilling program in lakes, lake bed outcrops, and ocean basins surrounding the regions where hominins evolved and a major investment in climate modeling experiments for key time intervals and regions that are critical to understanding human evolution.

How were the tropical Americas formed? This ambitious volume draws on extensive, multidisciplinary research to develop new views of the geological formation of the isthmus linking North and South America and of the major environmental changes that reshaped the Neotropics to create its present-day marine and terrestrial ecosystems. Recent discoveries show that dramatic changes in climate and ocean circulation can occur very quickly, and that ecological communities respond just as rapidly. Abrupt changes in the composition of fossil assemblages, formerly dismissed as artifacts of a poor fossil record, now are seen as accurate records of swift changes in the composition of ocean communities. The twenty-four contributors use current work in paleontology, geology, oceanography, anthropology, ecology, and evolution to paint this challenging portrait of rapid environmental and evolutionary change. Their conclusions argue for a revision of existing interpretations of the fossil record and the processes—including invading Eurasian peoples—that have produced it.

Most organisms and populations have to cope with hostile environments, threatening their existence. Their ability to respond phenotypically and genetically to these challenges and to evolve adaptive mechanisms is, therefore, crucial. The contributions to this book aim at understanding, from a evolutionary perspective, the impact of stress on biological systems. Scientists, applying different approaches spanning from the molecular and the protein level to individuals, populations and ecosystems, explore how organisms adapt to extreme environments, how stress changes genetic structure and affects life histories, how organisms cope with thermal stress through acclimation, and how environmental and genetic stress induce fluctuating asymmetry, shape selection pressure and cause extinction of populations. Finally, it discusses the role of stress in evolutionary change, from stress induced mutations and selection to speciation and evolution at the geological time scale. The book contains reviews and novel scientific results on the subject. It will be of interest to both researchers and graduate students and may serve as a text for graduate courses.