It is not often that one has the opportunity to send a public birthday greet ing to a friend and colleague of many years, and to congratulate him on having reached the age of reason. In fact it happens only once, and comes then as a surprise. Surely it was only a few years ago that we sat together at an International Genetics Congress in Ithaca, and only yesterday that we became members of the same department. The eighth floor of Schermerhorn Hall had a north end where the flies were and a south end furnished with mice, and in between, a seminar room and laboratory. There the distances were short and the doors open and the coffee pot busy. But it now appears that yesterday has fallen thirty years behind and that we have grown up. I find it interesting and appropriate that Dobzhansky's lifetime spans the period of maturation of the fields to which this volume is devoted. This is true in a chronological sense for his birth occurred in the same year, 1900, in which modern genetics began. The rediscovery of Mendel's princi ples and the interpretation of the nature of heredity and variation to which this event led were necessary prerequisites to the development of evolution ary biology as presented in this collection of essays.

Originally published in 2001, this is the second of two volumes published by Cambridge University Press in honour of Richard Lewontin. This second volume of essays honours the philosophical, historical and political dimensions of his work. It is fitting that the volume covers such a wide range of perspectives on modern biology, given the range of Lewontin's own contributions. He is not just a very successful practitioner of evolutionary genetics, but a rigorous critic of the practices of genetics and evolutionary biology and an articulate analyst of the social, political and economic contexts and consequences of genetic and evolutionary research. The volume begins with an essay by Lewontin on Natural History and Formalism in Evolutionary Genetics, and includes contributions by former students, post-docs, colleagues and collaborators, which cover issues ranging from the history and conceptual foundations of evolutionary biology and genetics, to the implications of human genetic diversity.