The Sipuncula, a group of ocean-dwelling worms related to annelids and mollusks, play a significant role in the bioerosion of coral reefs and are useful indicators of environmental conditions. The 155 species live in a wide variety of marine habitats at all depths, in sand and mud, in burrows in soft rock and dead coral, and inside such protective shelters as mollusk shells. Important food items for fish and invertebrate predators, they also recycle organic wastes and function as bioassay tools for human diseases such as cystic fibrosis and acute cholera. Edward B. Cutler brings together in this volume everything that is known about the entire phylum. An introduction, with practical information about collecting and handling the animals, is followed by Part One, which incorporates new systematic analyses made during the past twenty years and offers illustrated keys to all taxa, replacing the work of A.C. Stephen and S.J. Edmonds. Part Two reviews the past thirty years' work in such areas as ecology, muscular sysetms, blood chemistry, respiration, reproduction, and excretion. Part Three provides a new synthetic perspective on the phylum's zoogeography and evolutionary relationships, both to other phyla and within the phylum. It utilizes information from the fossil record, paleo-oceanographic data, and comparative studies of immunology, physiology, embryology, and anatomy.

The Sipuncula, a group of ocean-dwelling worms related to annelids and mollusks, play a significant role in the bioerosion of coral reefs and are useful indicators of environmental conditions. The 155 species live in a wide variety ofmarine habitats at all depths, in sand and mud, in burrows in soft rock and dead coral, and inside such protective shelters as mollusk shells. Important food items for fish and invertebrate predators, they also recycle organic wastes and function as bioassay tools for human diseases such as cystic fibrosis and acute cholera. Edward B. Cutler brings together in this volume everything that is known about the entire phylum.An introduction, with practical information about collecting and handling the animals, is followed by Part One, which incorporates new systematic analyses made during the past twenty years and offers illustrated keys to all taxa, replacing the work of A.C. Stephen and S.J. Edmonds. Part Two reviews the past thirty years' work in such areas as ecology, muscular sysetms, blood chemistry, respiration, reproduction, and excretion. Part Three provides a new synthetic perspective on the phylum's zoogeography and evolutionary relationships, both to other phyla and within the phylum. It utilizes information from the fossil record, paleo-oceanographic data, and comparative studies of immunology, physiology, embryology, and anatomy.Edward B. Cutler is Professor of Biology at Utica College of Syracuse University, now on long-term leave at the Museum of Comparative Zoology, Harvard University.

In June 1970, a diverse group of scientists attended the International Symposium on the Biology of the Sipuncula and Echiura at the Marine Biological Station in Kotor, Yugoslavia. Forty-two years later in June 2012, an aspiring generation of like-minded scientists convened the Second International Symposium on the Biology of the Sipuncula (ISBS2) along the banks of the Fort Pierce Inlet to the Indian River Lagoon in Florida, USA. The primary objective of the second symposium was to collect the world's dedicated sipunculan biologists in one place for a long overdue face-to-face communication of past, present and future research. We met our objective. This proceedings volume includes a brief summary of workshop discussions and field events, and a compilation of selected research papers presented by an international group of sixteen scientists from twelve nations in attendance at the ISBS2. Herein, we highlight molecular, developmental, morphological, ecological and biogeographic diversity of adult and larval sipunculans. And, we introduce several of the outstanding research challenges associated with resolving sipunculan interrelationships, establishing standard sets of taxonomic characters, refining methods for identification of cryptic species, reconstructing an evolutionary framework of developmental life history patterns, and addressing implications of recent phylogenetic and phylogenomic hypotheses that have relocated the ancient radiation of unsegmented sipunculan body plans within the predominantly segmented Annelida.--Provided by publisher.

Annelida, mainly consisting of marine Polychaeta and in faunal and partly parasitic Clitellata, is one the most significant metazoan taxa. Its more than 20.000 described species invade nearly all habitats and play a central role in marine benthic systems as well as in terrestrial soil communities. Annelids include all soft-bodied segmented worm-like organisms and have been recognized as a separate "phylum" for almost 200 years. Recently, evidence has been accumulated which shows that some of the groups formerly regarded as independent "phyla" such as Pogonophora (now recognized as Siboglinidae), Echiura, Myzostomida and perhaps Sipuncula, are most probably nothing else than greatly modified Annelida. The extreme morphological diversity found especially in Polychaeta displays the plasticity of a simple segmented organisation that basically is nothing else but a serial repetition of identical units. Thus, annelids are highly important to our understanding of fundamental questions about morphological and adaptive diversity, as well as clarifying evolutionary changes and phylogenetic relationships. The book aims to summarize our knowledge on Polychaetes polychaetes and their allies and gives an overview of recent advances gained by studies that employed conventional and modern methods plus, increasingly and importantly, the use of molecular markers and computer-assisted kinship analyses. It also reflects the state of art in polychaete sciences and presents new questions and controversies. As such it will significantly influence the direction of research on Polychaeta and their related taxa.

This book is the first in a series of 4 volumes in the Handbook of Zoology series about morphology, anatomy, reproduction, development, ecology, phylogeny and systematics of Annelida. This first volume covers members of the so-called basal radiation and the first part of Sedentaria. It is supplemented by chapters on the history of annelid research, their fossil record, and an introduction to the phylogeny of annelids and their position in the tree of life. In the latter chapter the history of their systematic is reviewed giving an almost complete picture of systematic-scientific progress especially in the past years which changed our view on annelid phylogeny dramatically. The most basal annelids, lately united as Palaeoannelida, represent two families of aberrant polychaetes formerly often suggested to be highly derived which now give us a fresh look on how the ancestral annelid may have looked like. These lack certain key characters such as nuchal organs and possess rather simple nervous systems which now likely represent primitive character states. In this basal radiation the first taxon of apparently unsegmented and achaetigerous animals is positioned, the Sipuncula. Most likely another group of platyhelminth-like and unsegmented and even chaeta-lees annelids, Lobatocerebridae falls into this basal radiation. The section of Sedentaria starts with Orbiniida, a taxon characterized by elongated, thread-like worms which do not have anterior appendages like palps and comprises several families representing members of the Meiofauna. These minute worms often inhabiting the interstitial spaces in marine sands are suggested to have evolved by progenesis. The second higher taxon is represented by Cirratuliformia comprising nine families of typical sedentary polychaetes each of which showing a remarkable variation of the annelid body plan. Members of this taxon usually exhibit many annelid characters but certain also lack the most typical prostomial appendages, the palps.